User blog comment:Golby Face!/Subspecies or Relative Species?/@comment-26434953-20171225161715

So by extension, the Hoopfish/Spinefish, Biter/Blighter, and Cave Crawler and Blood Crawler aren't subspecies?

The definition for subspecies is "a taxonomic category that ranks below species, usually a fairly permanent geographically islotated race". So by this definition, the Oculus would definitely fit as a subspecies to the Peeper. The PDA entry even supports this.